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​# = Lab graduate students, + = Lab undergraduate students

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  1. Wan, H. Y., M. A. Lommler, S. A. Cushman, J. S. Sanderlin, J. L. Ganey, A. J. Sánchez Meador, and P. Beier. 2025. A multi-level, multi-scale comparison of LiDAR-and LANDSAT-based habitat selection models of Mexican spotted owls in a post-fire landscape. Ecological Informatics 89:103168. https://doi.org/10.1016/j.ecoinf.2025.103168

  2. Bao, S., H. Zhu, P. Wester, W. Wang, Z. Li, B. Ren, D. Yan, and H. Y. Wan. 2025. Wild bees outperform managed bees in density-dependent pollination for alfalfa reproduction. Agriculture, Ecosystems and Environment 389:109693. https://doi.org/10.1016/j.agee.2025.109693

  3. Li, Y., Z. Xia, L. Nguyen, H. Y. Wan, L. Wan, M. Wang, N. Jia, V. R. R. Matli, Y. Li, M. Seeley, E. F. Moran, and J. Liu. 2025. Spatiotemporal dynamics of coastal dead zones in the Gulf of Mexico over 20 years using remote sensing. Science of The Total Environment 979:179461. https://doi.org/10.1016/j.scitotenv.2025.179461

  4. #Hysen, L., H. Y. Wan, P. Jantz, J. Gagnon, and S. A. Cushman. 2025. Variability in habitat selection between herds for a widespread ungulate. Ecological Modelling 501:110991. https://doi.org/10.1016/j.ecolmodel.2024.110991

  5. Lichwa-Schneringer, E., J. W. Cain III, H. Y. Wan, G. Fuller, C. Millberry, and M. S. Gunther. 2025. Ecological and social drivers of Mexican wolf home range size across spatiotemporal scales. Journal of Mammalogy 106(1):105-117. https://doi.org/10.1093/jmammal/gyae110

  6. #Becdach, R. E., +K. Larson, +K. Crouch, E. Meisman, A. Goldman, C. L. Chambers, M. DeJong, V. Titus, M S. Gunther, and H. Y. Wan. 2024. A celebration and reflection on the equity trend between women and men in wildlife publishing. Journal of Wildlife Management 89(1):e22680. https://doi.org/10.1002/jwmg.22680

  7. Martínez-Fonseca, J. G., E. P. Westeen, H. Y. Wan, S. A. Cushman, and C. L. Chambers. 2024. A global review of landscape-scale analyses in bats reveals geographic and taxonomic biases and opportunities for novel research. Biological Conservation 299:110829. https://doi.org/10.1016/j.biocon.2024.110829

  8. Feng, Y., K. Zhang, A. Li, Y. Zhang, K. Wang, N. Guo, H. Y. Wan, X. Tan, N. Dong, X. Xu, R. He, B. Wang, L. Fan, S. Ge, and P. Song. 2024. Spatial and seasonal variation and the driving mechanism of the thermal effects of urban park green spaces in Zhengzhou, China. Land 13(9):1474. https://doi.org/10.3390/land13091474

  9. #Nayeri, D., S. Cushman, J. Ganey, #L. Hysen, M. S. Gunther, D. Willey, and H. Y. Wan. 2024. Multiscale habitat suitability modeling for a threatened raptor offers insight into ecological model transferability. Ecological Modelling 496:110845. https://doi.org/10.1016/j.ecolmodel.2024.110845

  10. Zanuari, A. H., K. Z. Abidin, M. S. Mansor, H. Y. Wan, S. N. A. S. Abdullah, P. Abdul-Patah, and S. M. Nor. 2024. Identifying priority corridors and bottlenecks for three threatened large mammal species in the oil palm-dominated landscape of Peninsular Malaysia. Global Ecology and Conservation 54:e03092. https://doi.org/10.1016/j.gecco.2024.e03092

  11. Eshtiaghi, A., S. Naderi, A. Mohammadi, and H. Y. Wan. 2024. Identifying wild boar (Sus scrofa) crop damage hotspots to mitigate human-wild boar conflicts in northern Iran. Global Ecology and Conservation 54:e03065. https://doi.org/10.1016/j.gecco.2024.e03065

  12. Pan, G., H. Y. Wan, D. R. Nash, K. Shi, and S. Cushman. 2024. Snow leopards exhibit non-stationarity in scale-dependent habitat selection between two national protected areas in China. Ecological Modelling 494:110759. https://doi.org/10.1016/j.ecolmodel.2024.110759

  13. Zhang, Y., Z. Ma, X. Li, J. Wu, Z. Li, Z. Yang, H. Y. Wan, D. Pan, S. Jiang, X. Yue, Y. Li, D. Wang, and J. Liu. 2024. Soil phosphorus availability is enhanced by nitrogen and litter addition during the growing season. Plant and Soil. https://doi.org/10.1007/s11104-024-06660-4

  14. Sartor, C. C., S. A. Cushman, H. Y. Wan, F. P. Tirelli, S. L. Althoff, E. Eizirik, T. C. Trigo, and T. R. O. de Freitas. 2024. Prioritizing conservation areas to mitigate connectivity loss and local extinction risk of a small carnivore (Leopardus guttulus) in South America. Biodiversity and Conservation 33:1605-1622. https://doi.org/10.1007/s10531-024-02817-w

  15. Cushman, S. A., K. Craighead, M. Yacelga, H. Y. Wan, Z. Kaszta, and D. W. Macdonald. 2023. Seventy-two models of large mammal connectivity across Panama: Insights into a critical biogeographic linkage zone. Frontiers in Ecology and Evolution 11:1250255. https://doi.org/10.3389/fevo.2023.1250255

  16. Feizabadi, H. A., A. Mohammadi, G. Shahnaseri, and H. Y. Wan. 2023. Comparing drivers and protection of core habitat and connectivity for two sympatric desert carnivores. Global Ecology and Conservation 48:e02696. https://doi.org/10.1016/j.gecco.2023.e02696

  17. Rezaei, S., A. Mohammadi, S. Shadloo, M. Ranaie, and H. Y. Wan. 2023. Climate change induces habitat shifts and overlaps among carnivores in an arid and semi-arid ecosystem. Ecological Informatics 77:102247. https://doi.org/10.1016/j.ecoinf.2023.102247

  18. Hassan, N., Z. Zhong, D. Wang, Y. Zhu, I. Naeen, A. B. Ahungu, H. Y. Wan, and X. Li. 2023. Effects of long-term mowing on species diversity, biomass, and composition of plant community in a semi-arid grassland in northeastern China. Applied Vegetation Science 26(3):e12743. https://doi.org/10.1111/avsc.12743

  19. Wang, Z., Z. Zhong, J. Cahill, E. Holden, H. Y. Wan, #L. Hysen, Y. Gao, Z. Wang, O. Batlai, J. Cushman, D. Wang, and X. Li. 2023. Standing litter modifies top-down effects of large herbivores on a grassland plant community. Ecosystems 26:1784-1795. https://doi.org/10.1007/s10021-023-00864-y

  20. #Hysen, L. B., S. A. Cushman, F. A. Fogarty, E. C. Kelly, #D. Nayeri, and H. Y. Wan. 2023. Northern spotted owl nesting habitat under high potential wildfire threats along the California Coastal Redwood Forest. Science of The Total Environment 890:163414. https://doi.org/10.1016/j.scitotenv.2023.163414

  21. Sun, Y., X. Zhu, L. Sui, Z. Zhu, B. Ren, D. Pan, H. Y. Wan, and H. Zhu. 2023. Effects of water stress on plant volatile emission and insect oviposition preference in an agroecosystem. Agricultural and Forest Entomology 25(3):458-467. https://doi.org/10.1111/afe.12567

  22. Dar, S. A., S. K. Singh, H. Y. Wan, S. A. Cushman, T. Bashir, and S. Sathyakumar. 2023. Future land use and climate change escalate connectivity loss for Himalayan brown bears. Animal Conservation 26(2):199-215. https://doi.org/10.1111/acv.12813

  23. Khosravi, R., H. Y. Wan, M.-R. Sadeghi, and S. A. Cushman. 2023. Identifying human-brown bear conflict hotspots for prioritizing critical habitat and corridor conservation in southwestern Iran. Animal Conservation 26(1):31-45. https://doi.org/10.1111/acv.12800

  24. Shirk, A. J., G. M. Jones, Z. Yang, R. J. Davis, J. L. Ganey, R. J. Gutiérrez, S. P. Healey, S. J. Hedwall, S. J. Hoagland, R. Maes, K. Malcolm, K. S. McKelvey, C. Vynne, J. S. Sanderlin, M. K. Schwartz, M. E. Seamans, H. Y. Wan, and S. A. Cushman. 2023. Automated habitat monitoring systems linked to adaptive management: a new paradigm for species conservation in an era of rapid environmental change. Landscape Ecology 38:7-22. https://doi.org/10.1007/s10980-022-01457-1

  25. Jones, G. M., A. J. Shirk, Z. Yang, R. J. Davis, J. L. Ganey, R. J. Gutiérrez, S. P. Healey, S. J. Hedwall, S. J. Hoagland, R. Maes, K. Malcolm, K. S. McKelvey, J. S. Sanderlin, M. K. Schwartz, M. E. Seamans, H. Y. Wan, and S. A. Cushman. 2023. Spatial and temporal dynamics of Mexican spotted owl habitat in the southwestern US. Landscape Ecology 38:23–37. https://doi.org/10.1007/s10980-022-01418-8

  26. #Hysen, L., #D. Nayeri, S. A. Cushman, and H. Y. Wan. 2022. Background sampling for multi-scale ensemble habitat selection modeling: does the number of points matter? Ecological Informatics 72(2022):101914. https://doi.org/10.1016/j.ecoinf.2022.101914

  27. Sartor, C. C., H. Y. Wan, J. A. Pereira, E. Eizirik, T. C. Trigo, T. R. O. de Freitas, and S. A. Cushman. 2022. Landscape genetics outperforms habitat suitability in predicting landscape resistance for congeneric cat species. Journal of Biogeography 49(12):2206-2217. https://doi.org/10.1111/jbi.14498

  28. #Nayeri, D., A. Mohammadi, #L. Hysen, D. Hipólito, D. Huber, and H. Y. Wan. 2022. Identifying human-caused mortality hotspots to inform human-wildlife conflict mitigation. Global Ecology and Conservation 38(2022): e02241. https://doi.org/10.1016/j.gecco.2022.e02241

  29. Wang, G., S. A. Cushman, H. Y. Wan, M. Liu, and S. Jombach. 2022. Comparison of least-cost path and UNICOR cumulative resistant kernel analyses in mapping ecological connectivity networks in Luohe region, China. Journal of Digital Landscape Architecture 7:176-190. https://doi.org/10.14627/537724018

  30. Khosravi, R., H. R. Pourghasemi, R. Adavoudi, L. Julaie, and H. Y. Wan. 2022. A spatially-explicit analytical framework to assess wildfire risks on brown bear habitat and corridors in protected areas. Fire Ecology 18:1. https://doi.org/10.1186/s42408-021-00125-0

  31. Mohammadi, A., K. Almasieh, #D. Nayeri, M. A. Adibi, and H. Y. Wan. 2022. Comparison of habitat suitability and connectivity modelling for three carnivores of conservation concern in an Iranian montane landscape. Landscape Ecology 37:411–430. https://doi.org/10.1007/s10980-021-01386-5

  32. Craighead, K., M. Yacelga, H. Y. Wan, R. Vogt, and S. A. Cushman. 2022. Scale-dependent seasonal habitat selection by jaguars (Panthera onca) and pumas (Puma concolor) in Panama. Landscape Ecology 37:129–146. https://doi.org/10.1007/s10980-021-01335-2

  33. López-Márquez, V., S. A. Cushman, J. Templado, H. Y. Wan, H. M. Bothwell, and A. Machordom. 2021. Genetic connectivity of two marine gastropods in the Mediterranean Sea: seascape genetics reveals species-specific oceanographic drivers of gene flow. Molecular Ecology 30(19):4608-4629. https://doi.org/10.1111/mec.16080

  34. Zeller, K. A., C. A. Schroeder, H. Y. Wan, G. Collins, K. Denryter, A. F. Jakes, and S. A. Cushman. 2021. Forecasting habitat and connectivity for pronghorn across the Great Basin ecoregion. Diversity and Distributions 27(12):2315-2329. https://doi.org/10.1111/ddi.13402

  35. Dar, S. A., S. K. Singh, H. Y. Wan, V. Kumar, S. A. Cushman, and S. Sathyakumar. 2021. Projected climate change threatens Himalayan brown bear habitat more than human land use. Animal Conservation 24(4):659-676. https://doi.org/10.1111/acv.12671

  36. Mohammadi, A., K. Almasieh, H. Y. Wan, #D. Nayeri, A. Alambeigi, J. Ransom, and S. A. Cushman. 2021. Integrating Spatial Analysis and Questionnaire Survey to Better Understand Human-Onager Conflict in Southern Iran. Scientific Reports 11:12423. https://doi.org/10.1038/s41598-021-91921-w

  37. Lucid, M., H. Y. Wan, S. Ehlers, L. Robinson, L. Svancara, A. Shirk, and S. Cushman. 2021. Land Snail Microclimate Niches Identify Suitable Areas for Climate Refugia Management on a Montane Landscape. Ecological Indicators 129:107885. https://doi.org/10.1016/j.ecolind.2021.107885

  38. Chiaverini, L., H. Y. Wan, B. Hahn, A. Cilimburg, T. N. Wasserman, and S. A. Cushman. 2021. Effects of non-representative sampling design on multi-scale habitat models: flammulated owls in the Rocky Mountains. Ecological Modelling 450:109566. https://doi.org/10.1016/j.ecolmodel.2021.109566

  39. Sartor, C., S. A. Cushman, H. Y. Wan, R. Kretschmer, J. A. Pereira, N. Bou, M. Cosse, S. González, E. Eizirik, T. R. O. de Freitas, and T. C. Trigo. 2021. The role of the environment in the spatial dynamics of an extensive hybrid zone between two Neotropical cats. Journal of Evolutionary Biology 34 (4):614-627. https://doi.org/10.1111/jeb.13761

  40. Bosco, L., S. A. Cushman, H. Y. Wan, K. A. Zeller, R. Arlettaz, and A. Jacot. 2021. Fragmentation effects on woodlark habitat selection depend on habitat amount and spatial scale. Animal Conservation 24(1):84-94. https://doi.org/10.1111/acv.12604

  41. Li, X., S. Wang, C. Prather, H. Y. Wan, H. Zhu, P. Nummi, M. Inbar, Q. Gao, D. Wang, and Z. Zhong. 2021. Large herbivores facilitate an insect herbivore by modifying plant community composition in a temperate grassland. Ecology and Evolution 11(22):16314-16326. https://doi.org/10.1002/ece3.8327

  42. Wang, G., S. A. Cushman, H. Y. Wan, H. Li, Z. Szabó, D. Ning, and S. Jombach. 2021. Ecological Connectivity Networks for Multi-dispersal Scenarios Using UNICOR Analysis in Luohe Region, China. Journal of Digital Landscape Architecture 6:230-244. http://doi.org/10.14627/537705020

  43. Wan, H. Y., S. A. Cushman, and J. L. Ganey. 2020. The effect of scale in quantifying fire impacts on species habitats. Fire Ecology 16:9. https://doi.org/10.1186/s42408-020-0068-2

  44. Ashrafzadeh, M. R., R. Khosravi, M. A. Adibi, A. T. Tehrani, H. Y. Wan, and S. A. Cushman. 2020. A multi-scale, multi-species approach for assessing effectiveness of habitat and connectivity conservation for endangered felids. Biological Conservation 245:108523. http://doi.org/10.1016/j.biocon.2020.108523

  45. Jones, G. M., R. J. Gutiérrez, W. M. Block, P. C. Carlson, E. J. Comfort, S. A. Cushman, R. J. Davis, S. A. Eyes, A. B. Franklin, J. L. Ganey, S. Hedwall, J. J. Keane, R. Kelsey, D. Lesmeister, M. P. North, S. L. Roberts, J. T. Rockweit, J. S. Sanderlin, S. C. Sawyer, B. Solvesky, D. J. Tempel, H. Y. Wan, A. L. Westerling, G. C. White, and M. Z. Peery. 2020. Spotted owls and forest fire: Comment. Ecosphere 11(12):e03312. https://doi.org/10.1002/ecs2.3312  

  46. Wan, H. Y., S. A. Cushman, and J. L. Ganey. 2019. Improving habitat and connectivity model predictions with multi-scale resource selection functions from two geographic areas. Landscape Ecology 34(3):503-519. https://doi.org/10.1007/s10980-019-00788-w

  47. Wan, H. Y., S. A. Cushman, and J. L. Ganey. 2019. Recent and projected future wildfire trends across the ranges of three spotted owl subspecies under climate change. Frontiers in Ecology and Evolution 7:37. https://doi.org/10.3389/fevo.2019.00037

  48. Bosco, L., H. Y. Wan, S. A. Cushman, R. Arlettaz, and A. Jacot. 2019. Separating the effects of habitat amount and fragmentation on invertebrate abundance using a multi-scale framework. Landscape Ecology 34(1):105-117. https://doi.org/10.1007/s10980-018-0748-3

  49. López-Márquez, V., S. A. Cushman, J. Templado, H. Y. Wan, H. Bothwell, C. Kruschel, V Macic, and A. Machordom. 2019. Seascape genetics and connectivity modelling for an endangered Mediterranean coral in the Adriatic Sea. Landscape Ecology 34(11):2649–2668. https://doi.org/10.1007/s10980-019-00911-x

  50. Kapsar, K. E., C. Hovis, R. F. B. da Silva, E. K. Buchholtz, A. K. Carlson, Y. Dou, Y. Du, P. R. Furumo, Y. Li, A. Torres, D. Yang, H. Y. Wan, J. G. Zaehringer, and J. Liu. 2019. Telecoupling research: the first five years. Sustainability 11(4):1033. https://doi.org/10.3390/su11041033

  51. Yang, D., H. Y. Wan, T. Huang, and J. Liu. 2019. The role of citizen science in conservation under the telecoupling framework. Sustainability 11(4):1108. https://doi.org/10.3390/su11041108

  52. Wan, H. Y., J. L. Ganey, and S. A. Cushman. 2018. Habitat fragmentation reduces genetic diversity and connectivity of the Mexican spotted owl: a simulation study using empirical resistance models. Genes 9(8):403. https://doi.org/10.3390/genes9080403

  53. Wan, H. Y., J. L. Ganey, C. D. Vojta, and S. A. Cushman. 2018. Managing emerging threats to spotted owls. Journal of Wildlife Management 82(4):682-697. https://doi.org/10.1002/jwmg.21423

  54. Littell, J. S., D. McKenzie, H. Y. Wan, and S. A. Cushman. 2018. Climate change and future wildfire in the western United States: an ecological approach to nonstationarity. Earth's Future 6(8):1097-1111. https://doi.org/10.1029/2018EF000878

  55. Wan, H. Y., K. McGarigal, J. L. Ganey, V. Lauret, B. C. Timm, and S. A. Cushman. 2017. Meta-replication reveals non-stationarity in multi-scale habitat selection of Mexican spotted owl. Condor 119(4):641-658. https://doi.org/10.1650/CONDOR-17-32.1

  56. Ganey, S. A., H. Y. Wan, S. A. Cushman, and C. D. Vojta. 2017. Conflicting perspectives on spotted owls, wildfire, and forest restoration. Fire Ecology 13(3):146-165. https://doi.org/10.4996/fireecology.130318020

  57. Rhodes, A. C, H. Y. Wan, and S. B. St. Clair. 2017. Herbivory impacts of elk, deer and cattle on aspen forest recruitment along gradients of stand composition, topography and climate. Forest Ecology and Management 397:39-47. https://doi.org/10.1016/j.foreco.2017.04.014

  58. McGarigal, K., H. Y. Wan, K. A. Zeller, B. C. Timm, and S. A. Cushman. 2016. Multi-scale habitat modeling: a review and outlook. Landscape Ecology 31(6):1161-1175. https://doi.org/10.1007/s10980-016-0374-x

  59. Wan, H. Y., A. C. Rhodes, and S. B. St. Clair. 2014. Fire severity alters plant regeneration patterns and defense against herbivores in mixed aspen forests. Oikos 123:1479-1488. https://doi.org/10.1111/oik.01521

  60. Wan, H. Y., A. C. Olson, K. D. Muncey, and S. B. St. Clair. 2014. Legacy effects of fire size and severity on forest regeneration, recruitment, and wildlife activity in aspen forests. Forest Ecology and Management 329:59-68. https://doi.org/10.1016/j.foreco.2014.06.006

©2025 by Ho Yi Wan

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